The protests on Sunday were directed at the government and Prime Minister Andrej Babis' plans to change how public service broadcasters are funded - replacing licence fees with state budget funding and making accompanying cuts.
Mikulas Minar, from the organization Million Moments for Democracy, said between 350,000 and 450,000 protesters had gathered on Sunday afternoon. Police and the authorities have not commented on the number of protesters.
Babis' coalition government, which includes the far-right SPD party, claims that the funding changes will affect neither the independence nor the content of public service broadcasters.
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